O3,CO2浓度及太阳光谱变化对作物光合作用影响的数值模拟研究
NUMERICAL SIMULATION OF THE INFLUENCE OF O3,CO2 AND SPECTRUM VARIATION ON THE PHOTOSYNTHESIS OF CROP CANOPY
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摘要: 对光合作用蒸腾作用气孔调节进行耦合,从生物化学尺度扩展至冠层尺度,发展了一个冬小麦冠层光合作用生态动力模式,模式考虑了O3,CO2和光谱变化对作物光合的综合影响。利用美国光合作用实测资料对模式进行验证,叶片模式通过了相关显著性检验并具有较高的准确度。数值分析表明:当O3浓度由0×109V/V上升至200×109V/V时,冠层光合速率下降29%左右;当CO2浓度由330×106V/V上升至660×106V/V时,冠层光合速率增加大约37%;当光谱比例系数由目前的0.5下降至0.4时,冠层光合速率将下降27%左右。对于污染严重、易发生光化学烟雾的城郊附近,在阳光强烈的典型晴天,中午O3浓度达到200×109V/V时,即使气候条件不发生改变,CO2浓度对作物光合作用的正效应也不足以弥补O3浓度升高所造成的负效应,冠层光合速率将比目前干洁地区略有下降,如果进一步考虑光合作用有效辐射光谱成分下降至0.4左右,冠层光合作用将比目前的BASE值下降35%左右。Abstract: Aleaf model was est ablished for simulating phot osynthesis of winter wheat based on charact eristics of physiological ecology of plant,and then a canopy photosy nthesis model was scrolled up from the leafmodel.The influence of CO2,O3 and spectrum variation on canopy photosynthesis was fully considered in the model.Validation of the leafmodel showed that the model could simulate photosynthesis rate of winter wheat in field fairly well.The simulation result by using the model indicated that:(a) When the concent ration of O3 increased from 0×10-9V/V to 200×10-9V/V,the canopy photosynthesis will decrease about 29% in contrast with the base value.The canopy photosynthesis will enhance about 37% when the concentration of CO2 changed from 330×10-6V/V to 660×10-6V/V.The canopy photosynthesis declined with the debasement of spectrum coefficient according to the linear relation,which reduced about 27% with the spectrum coefficient varied from 0.5 to 0.4.(b) The canopy photosynthesis will be reduced about 35% in suburb,when the spectrum coefficient reduced to 0.4 and the concentration of O3 increased to 200×10-9V/V,even with a doubled CO2 concentration and unchanged climatic scenarios.